§16 · parent-of-origin

Parent-of-origin is a duration asymmetry: sustained maternal flips, transient paternal reverts

At equal supra-spinodal amplitude (0.8759) the crossing is set by the duration of exposure: a sustained maternal drive flips a held switch (end state 1.4361), while a transient paternal burst of the same amplitude relaxes back (-1.2052). The inherited sign is the maternal sign, and the asymmetry holds at every measured germline locus (13/13). [V]/[O].

On the deep germline master REC8 (γ = 1.4525, spinodal 0.6738) a held switch needs 342 steps of supra-spinodal exposure to cross. The sustained maternal context supplies it and the switch flips; a transient paternal burst of 119 steps (decay τ = 102) relaxes back — by duration, not amplitude. Under opposing parents the maternal sign is inherited and agreement reinforces (94 vs 342 steps).

Same amplitude, different duration — the crossing is set by time

Sperm and egg deliver the same kind of object — a drive written at an A4 coordinate — in two temporal contexts. On REC8 (γ = 1.4525, spinodal 0.6738) both parents supply the same supra-spinodal amplitude (0.8759) — the fairness condition that keeps this read direction-only. A held switch needs 342 steps above the spinodal to cross. The egg's large cytoplasm holds the maternal payload sustained, so the switch flips (end state 1.4361); the sperm's small bolus is diluted to a transient burst (119 steps), which relaxes back to -1.2052. The asymmetry is in duration, not amplitude.

The inherited sign is the maternal sign

Under opposing parents, the dominant maternal sign is inherited: a maternal + against a paternal − settles ON (1.2052), and reversing the roles inherits OFF (-1.2052). When the two parents agree, the switch crosses faster — 94 steps versus 342 for the maternal context alone. The sign is read; the magnitude is not.

The asymmetry is universal across the germline atlas

The sustained-vs-transient asymmetry is not a property of one locus: it holds at 13 of 13 measured germline switches — every one flips under the sustained maternal context and reverts under the matched transient paternal burst.

Honest course-correction. PO3 was first hypothesised as “the paternal disadvantage rises with γ.” The substrate returned the opposite ordering: under equal-relative amplitude the descriptive rank-correlation between γ and crossing time came out ρ = -1.00 (crossing time is flat / slightly decreasing in γ). Per the inheritance discipline the claim was rewritten to the robust result — the asymmetry is universal in presence, not monotone in magnitude — with ρ reported as it falls, no directional claim graded, and no γ tuned. The absolute parent-of-origin penetrance stays [O].