Aging & Senescence · §2

Node atlas: measured promoter gamma

The four aging masters carry measured promoter γ: TP53 1.4298, CDKN2A 1.4424, TERT 1.5539, FOXO3 1.5942 — each computed as −mean SantaLucia nearest-neighbour ΔG37 over the TSS−2000..+500 window and never fitted. Emergence order by ascending γ is cellular senescence, arrest switch, telomere maintenance, longevity signaling.

γ is the promoter stacking-energy identity from DNA Layer 1; it fixes each node's barrier γ²/4 and dwell γ1.5. The four values are vendored from NCBI RefSeq accessions and reproduce bit-for-bit from the committed promoter cache.

The masters and their identity

Each aging master is a real promoter, and its γ is a measurement, not a parameter. The window is fixed (TSS−2000 to +500, 2501 bp), the nearest-neighbour table is SantaLucia 1998, and the value is −mean ΔG37 over the window. The same pipeline produces the DNA gene-clock γ.

The four measured aging masters. γ = −mean SantaLucia 1998 NN ΔG37 over TSS−2000..+500, vendored from NCBI RefSeq; never fitted.
GeneNodeγAccessionStrandDynamics
TP53cellular senescence1.429832NC_000017.11-stuck-attractor
CDKN2Asenescence arrest switch1.442444NC_000009.12-stuck-attractor
TERTtelomere maintenance1.553876NC_000005.10-reservoir-depletion
FOXO3longevity signaling1.594156NC_000006.12+maintenance

Why the order matters

γ sets two derived quantities. The barrier γ²/4 fixes how hard it is to leave a basin (state stability), and the dwell γ1.5 fixes how long a switch runs (reservoir capacity, §5). Sorting the masters by γ gives a fixed emergence order: cellular senescence (TP53) and the arrest switch (CDKN2A) sit lowest, then telomere maintenance (TERT), then longevity signaling (FOXO3).

γTP53 = 1.429832  |  barrier = γ²/4 = 0.5111

The diffuse node

The fifth node, systemic setpoint drift, has no single master gene. It is a circuit-level, derived quantity: the aggregate decline of every defended setpoint imported from the other packages. It is the object that §3 and §8 quantify.