§5 · transgenerational transmission

A parent's environment reaches a child only as a drive that survives two erasures

The SET is byte-identical parent→child; only the drive is inheritable. Two genome-wide reprogramming erasures square a mark's survival (p → p²), so heritability ranks by barrier — ascending γ, Spearman ρ = 0.995. The inherited sign is preserved, an RNA-only drive fades, a deep-barrier mark persists. [V].

At reprogramming noise D = 0.22, a shallow switch (γ = 1.34) survives one erasure with p = 0.6685 and inherits across two at p² = 0.4469, while a deep switch (γ = 1.4525) inherits at 0.5238 — deeper inherits more.

The genome itself does not move

Parent and child read the identical γ (DAZL = 1.3803, byte-identical); the environment never rewrites the SET. Whatever crosses to the next generation crosses as a drive on an unchanged switch.

The firewall squares survival

A germline mark must survive two genome-wide reprogramming erasures — in the primordial germ cells and again in the zygote — so its probability of reaching the next generation is multiplied, not added. Depth wins: at the same noise the deep switch inherits at p² = 0.5238 against the shallow switch's 0.4469.

Heritability is barrier ordering is γ ordering

Across the measured 13-gene germline atlas, survival rises monotonically with γ (Spearman ρ = 0.995). This is the central prediction: which marks inherit is set by the barrier, and the barrier is set by the measured promoter.

TG3 — germline heritability ranks ascending-γ (D = 0.22, 13 genes)
geneγp(survive one erasure)
TEKT11.34000.6685
NLRP51.34750.6740
MOS1.36190.6833
DAZL1.38030.6887
CATSPER11.40190.6983
DMC11.40560.7000
MLH11.40850.7015
SPO111.41050.7013
PRDM91.41650.7043
ZAR11.42260.7083
DNAH11.44690.7192
ZP31.44860.7208
REC81.45250.7238

Two carriers, two decay laws

The inherited sign is preserved through the firewall (ON stays ON, OFF stays OFF). But the two writers decay differently: an RNA-only drive dilutes each generation and drops below detectability by F3, whereas a deep-barrier methylation mark is still detectable across the same horizon. The generation index here is illustrative; the wild count is [O].

TG5 — decay ORDERING: RNA dilutes (rate 0.45), methylation retains (0.7238)
generationRNA amplitudemethylation amplitude
F01.00001.0000
F10.45000.7238
F20.20250.5238
F30.09110.3791
F40.04100.2744
F50.01850.1986
F60.00830.1437
FIREWALL · WHICH switch, the inherited SIGN, and the heritability ORDERING / DECAY are read; the absolute inherited phenotype magnitude and the wild generation-count are runtime [O]. Every clinical application — vaccination, gene therapy, fertility care — is handed to clinicians and regulators. see the [O] ledger →