§5 · transgenerational transmission
A parent's environment reaches a child only as a drive that survives two erasures
The SET is byte-identical parent→child; only the drive is inheritable. Two genome-wide reprogramming erasures square a mark's survival (p → p²), so heritability ranks by barrier — ascending γ, Spearman ρ = 0.995. The inherited sign is preserved, an RNA-only drive fades, a deep-barrier mark persists. [V].
At reprogramming noise D = 0.22, a shallow switch (γ = 1.34) survives one erasure with p = 0.6685 and inherits across two at p² = 0.4469, while a deep switch (γ = 1.4525) inherits at 0.5238 — deeper inherits more.
The genome itself does not move
Parent and child read the identical γ (DAZL = 1.3803, byte-identical); the environment never rewrites the SET. Whatever crosses to the next generation crosses as a drive on an unchanged switch.
The firewall squares survival
A germline mark must survive two genome-wide reprogramming erasures — in the primordial germ cells and again in the zygote — so its probability of reaching the next generation is multiplied, not added. Depth wins: at the same noise the deep switch inherits at p² = 0.5238 against the shallow switch's 0.4469.
Heritability is barrier ordering is γ ordering
Across the measured 13-gene germline atlas, survival rises monotonically with γ (Spearman ρ = 0.995). This is the central prediction: which marks inherit is set by the barrier, and the barrier is set by the measured promoter.
| gene | γ | p(survive one erasure) |
|---|---|---|
| TEKT1 | 1.3400 | 0.6685 |
| NLRP5 | 1.3475 | 0.6740 |
| MOS | 1.3619 | 0.6833 |
| DAZL | 1.3803 | 0.6887 |
| CATSPER1 | 1.4019 | 0.6983 |
| DMC1 | 1.4056 | 0.7000 |
| MLH1 | 1.4085 | 0.7015 |
| SPO11 | 1.4105 | 0.7013 |
| PRDM9 | 1.4165 | 0.7043 |
| ZAR1 | 1.4226 | 0.7083 |
| DNAH1 | 1.4469 | 0.7192 |
| ZP3 | 1.4486 | 0.7208 |
| REC8 | 1.4525 | 0.7238 |
Two carriers, two decay laws
The inherited sign is preserved through the firewall (ON stays ON, OFF stays OFF). But the two writers decay differently: an RNA-only drive dilutes each generation and drops below detectability by F3, whereas a deep-barrier methylation mark is still detectable across the same horizon. The generation index here is illustrative; the wild count is [O].
| generation | RNA amplitude | methylation amplitude |
|---|---|---|
| F0 | 1.0000 | 1.0000 |
| F1 | 0.4500 | 0.7238 |
| F2 | 0.2025 | 0.5238 |
| F3 | 0.0911 | 0.3791 |
| F4 | 0.0410 | 0.2744 |
| F5 | 0.0185 | 0.1986 |
| F6 | 0.0083 | 0.1437 |